Alan Covich

Professor Emeritus
Graduate Program Faculty

Professional Website


Ph.D. – Yale University

Areas of Expertise
Honors, Awards, and Achievements
  • Fellow, Society for Freshwater Science – 2019
  • Distinguished Ecologist, Colorado State University – 2018
  • Distinguished Service Award, Ecological Society of America- 2015
  • Fellow, Ecological Society of America- 2012
  • President, International Association for Ecology (INTECOL) – 2009
  • President, Ecological Society of America. – 2007
  • Fulbright Scholar, University of Coimbra, Portugal – 2003
  • Visiting Scholar, National Center for Ecological Synthesis and Analysis – 2001
  • Fellow, American Association for Advancement of Science – 2000
  • President, American Institute of Biological Sciences – 1999
  • American Water Resources Association Award of Excellence – 1997
  • President, North American Benthological Society – 1996

Professional Society Affiliations

  • American Association for the Advancement of Science
  • American Institute of Biological Sciences
  • Association for the Science of Limnology and Oceanography
  • Association for Tropical Biology and Conservation
  • International Association for Ecology
  • International Society for Limnology
  • Society for Freshwater Science
Courses Taught

ECOL 3400, ECOL 3500, ECOL 4310/6310, ECOL 8230

Research Interests
  • Impacts of floods and hurricanes on tropical stream food webs.
  • Drought impacts on terrestrial-freshwater linkages in temperate and tropical stream ecosystems.

Research Projects

Since 1988 I have worked with colleagues in Puerto Rico on the effects of urban growth, harvesting of freshwater shrimps and crabs, and intersections of road networks on river networks. I am also collaborating with Brazilian colleagues in Campo Grande, Mato Grosso do Sul on food webs in karst streams that flow into the Pantanal.

Selected Publications

Valente-Neto, F., F.H. da Silva, A.P. Covich, F. de Oliveira Roque. 2019. Streams dry and ecological uniqueness rise: Environmental selection drives aquatic insect patterns in a riverine network prone to intermittence. Hydrobiologia (Online:

Lininger, K.B., J. Raimondi, N. Kramer, D. Homrighausen, and A. Covich. 2019. Comparison of discharge pulses in temperate and tropical rainforest headwater stream networks. Journal of Hydrology 579:1242362.

de Oliveira Roque, F., F. Valente-Neto, M.E. Rodrigues, F. N. Souza, D.G. Nogueira, A. C. Piovezan-Borges, A.P. Covich and M. Samways. 2019. The Tinbergen shortfall: developments on aquatic insect behavior that are critical for freshwater conservation. Pp. 365-380, In: K. Del-Claro and R. Guillermo (eds.) Aquatic insects: Behaviour and Ecology, Springer, Cham.

Czaja, A., A.P. Covich, J.L. Estrada-Rodríguez, U. Romero-Méndez, J. Saenz-Mata, I.G. Meza-Sánchez, V. Ávila-Rodríguez, J. L. Becerra-López, D. R. Aguillón-Gutiérrez, and J.G. Castañeda-Gaytán. 2019. Freshwater snails from northern Mexico – A case of a “silent” local extinction with the description of a new species. Boletín de la Sociedad Geólogica Mexicana 71:609-629.

Corrêa, E.C., F. de Oliveira Roque, R.M. Utz; J. Sousa Corrêa, F. Leandro de Souza, and A.P. Covich. 2018. Effects of macroconsumers on benthic communities: rapid increases in dry-season accrual of calcium in a tropical karst stream. PLOS One 13: e0209102.

Shivers, S.D., S.W. Golladay, M.N. Waters, S.B. Wilde, P.D. Ashford, and A.P. Covich. 2018. Rivers to reservoirs: Hydrological drivers control reservoir function by affecting the abundance of submerged and floating macrophytes. Hydrobiologia 815:21-35

Marzolf, N., S. Golladay, P. McCormick, A. Covich, and S. Wilde. 2017. Inter- and intra-annual apple snail egg mass dynamics in a large southeastern US reservoir. Hydrobiologia 811:155-171.

Shivers, S.D., S.W. Golladay, M.N. Waters, S.B. Wilde, P.D. Ashford, and A.P. Covich. 2017. Changes in submerged aquatic vegetation (SAV) coverage caused by extended drought and flood pulses. Lake and Reservoir Management 34:218-229.

Smith, C.R., P.V. McCormick, A.P. Covich, and S.W. Golladay. 2017. Comparison of macroinvertebrate assemblages across a gradient of flow-permanence in an agricultural watershed. River Research and Applications 33:1428-38.

Covich, A.P. 2017. Frank Golley’s perspectives on environmental ethics and literacy: How to avoid irreversible impacts of hydro-power and inter-oceanic canal development on Meso-American tropical ecosystems? Human Ecology Review 23:39-53.

McKay, S.K, M.C. Freeman, and A.P. Covich. 2016. Application of effective discharge analysis to environmental flow decision making. Environmental Management 57:1153-1165.

Huete-Pérez, J.A., M. Ortega-Hegg, G.R. Urquhart, A.P. Covich, K. Vammen, B.E. Rittmann, J. C. Miranda, S. Espinoza-Corriols, A. Acevedo, M.L. Acosta, J.P. Gómez, M.T. Brett, M. Hanemann, A. Härer, J. Incer-Barquero, F.J. Joyce, J.W. Lauer, J.M. Maes, M.B. Tomson, A. Meyer, S. Montenegro-Guillén, W. L. Whitlow, J. L. Schnoor, P. J.J. Alvarez. 2016. Critical uncertainties and gaps in the Environmental and Social Impact Assessment of the proposed interoceanic canal through Nicaragua. BioScience 66:632-645.

Pérez-Reyes, O, T.A. Crowl, and A.P. Covich. 2015. Comparison of decapod communities across an urban-forest land use gradient in Puerto Rican streams. Urban Ecosystems. DOI:10.1007/s11252-015-00490-4.

Pérez-Reyes, O, T.A. Crowl, and A.P. Covich. 2015. Effects of elevation, food supplies, and water temperature on growth rates of freshwater topical shrimp in Luquillo Mountains, Puerto Rico. Freshwater Biology 60:1514-1524.

Patton, D., J.C. Bergstrom, R. Moore, and A.P. Covich. 2015. Economic value of carbon storage in US National Wildlife Refuge wetland ecosystems. Ecosystem Services 16:94-104.

Covich, A.P. 2015. Frank Golley: Interdisciplinary insights for a 21st century Earth stewardship, pp. 431-450, In: Earth stewardship: Linking ecology and ethics in theory and praxis, R. Rozzi, S.T.A. Pickett, J.B. Callicott, T. Chapin, J.J. Armesto and M.E, Power (eds.). Springer, New York.

Covich, A.P. 2015. Projects that never happened: Ecological insights from Darien, Panama. Bulletin of the Ecological Society of America.96:54-63.

Covich, A.P. 2014. Freshwater crustaceans: Adaptations to complex inland habitats and species interactions. Pp. 337-378, In: Natural History of Crustacea: Life styles and feeding biology of the Crustacea, Volume II. M. Thiel and L. Watling (eds.), Oxford University Press, Oxford.

Rincón, J., and A.P. Covich. 2014. Effects of insect and decapod exclusion and leaf litter species identity on breakdown rates in a tropical headwater stream. International Journal of Tropical Biology and Conservation 62: 143-154.

Thorp, J.H. and A.P. Covich. 2014. Overview and inland water habitats. Pp. 23-56. In: Thorp and Covich’s Freshwater Invertebrates. Volume I. Ecology and General Biology. Thorp, J.H. and D.C. Rogers (eds.). Academic Press, Elsevier, Boston.

Brokaw, N., J. Zimmerman, M. Willig, G. Camilo, A. Covich, T. Crowl, N. Fetcher, B. Haines, M. Gannon, J. Lodge, A. Lugo, R. Myster, C. Pringle, J. Sharpe, F. Scatena, T. Schowalter, W. Silver, J. Thompson, D. Vogt, K. Vogt, R. Waide, L. Walker, L. Woolbright, J. Wunderle, X. Zou. 2012. Response to Disturbance. Pp. 201-271 IN: N. Brokaw, T.A. Crowl, A.E. Lugo, W.H. McDowell, F.N. Scatena, R.H. Waide, and M.R. Willig (eds.), A Caribbean forest tapestry- The multi-dimensional nature of disturbance and response. Oxford University Press, Oxford, UK.

Crowl, T.A., N. Brokaw, R.B. Waide, G. González, K. Beard, E. Greathouse, A.E. Lugo, A.P. Covich, D. J. Lodge, C. Pringle, J. Thompson, and G.E. Belovsky. 2012. When and where biota matter: Linking disturbance regimes, species characteristics and dynamics of communities and ecosystems. pp. 272-304, IN: N. Brokaw, T.A. Crowl, A.E. Lugo, W.H. McDowell, F.N. Scatena, R.H. Waide, and M.R. Willig (eds.), A Caribbean forest tapestry- The multi-dimensional nature of disturbance and response. Oxford University Press, Oxford, UK.

Willig, M.R., C.P. Bloch, A.P. Covich, C. A. S. Hall, D. J. Lodge, A.E. Lugo, W.L. Silver, R. B. Waide, L.R. Walker, and J.K. Zimmerman. 2012. Long-term Research in the Luquillo Mountains: Synthesis and Foundations for the Future.
pp. 361-441, IN: N. Brokaw, T.A. Crowl, A.E. Lugo, W.H. McDowell, F.N. Scatena, R.H. Waide, and M.R. Willig (eds.), A Caribbean forest tapestry- The multi-dimensional nature of disturbance and response. Oxford University Press, Oxford, UK.

Hein, C.L., A.S. Pike, J.F. Blanco, A.P. Covich, F.N. Scatena, C.P. Hawkins, and T.A. Crowl. 2011. Effects of coupled natural and anthropogenic factors on the community structure of diadromous fish and shrimp species in tropical island streams. Freshwater Biology 56: 1002-1015.